†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
1









	




















































 


The
tables
in
this
appendix
summarize
the
life
history
parameter
values
used
by
EPA
to
calculate
age
1
equivalents,
fishery
yield,
and
production
foregone
from
I&
E
data
for
the
Ohio
River
case
study.

!

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

	


#




$

%


 



$
 





























&
Stage
Name
Natural
Mortality
(
per
stage)
a
Fishing
Mortality
(
per
stage)
b
Fraction
Vulnerable
to
Fisheryc
Weight
(
lbs)
d
Eggs
2.87
0
0
0.0000000202e
Larvae
1.72
0
0
0.00214f
Age
0
2.98
0
0
0.00851g
Age
1+
0.548
0
0
1.14g
Age
2+
0.548
0
0
1.82g
Age
3+
0.548
0
0
2.63g
Age
4+
0.548
0
0
3.48g
Age
5+
0.548
0
0
4.64g
Age
6+
0.548
0
0
5.04g
Age
7+
0.548
0
0
11.1g
Age
8+
0.548
0
0
12.7g
Age
9+
0.548
0
0
16.8g
Age
10+
0.548
0
0
27.8g
Age
11+
0.548
0
0
28g
Age
12+
0.548
0
0
36.1g
Age
13+
0.548
0
0
36.2f
Age
14+
0.548
0
0
36.3f
Age
15+
0.548
0
0
36.5f
a
Bartell
and
Campbell,
2000.
b
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
c
Assumed.
d
Weight
calculated
from
length
using
the
formula:
(
2.399*
10­
6)*
Length(
mm)
3.30
=
Weight(
gm)
(
Carlander,
1969).
e
Kleinholz,
2000.
f
Assumed
based
on
Carlander,
1969.
g
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
2
!

"



	


#
'


$



(

$
 






























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisherye
Weight
(
lbs)
f
Eggs
2.3a
0
0
0.000000559g
Larvae
4.61b
0
0
0.000186h
Age
0+
1.39b
0
0
0.00132i
Age
1+
0.446c
0
0
0.0362i
Age
2+
0.446c
0
0
0.0797i
Age
3+
0.446c
0
0
0.137i
Age
4+
0.446c
0
0
0.233i
Age
5+
0.446c
0
0
0.402i
Age
6+
0.446c
0
0
0.679i
Age
7+
0.446c
0
0
0.753i
Age
8+
0.446c
0
0
0.815i
Age
9+
0.446c
0
0
0.823j
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
survival
(
PSEG,
1999c)
for
channel
catfish
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
Calculated
from
survival
(
PSEG,
1999c)
for
brown
bullhead
(
Carlander,
1969)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Assumed.
f
Weight
calculated
from
length
using
the
formula:
(
8.797*
10­
6)*
Length(
mm)
3.060
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
g
Wang,
1986b.
h
Assumed
based
on
Wang,
1986b
and
Carlander,
1969.
i
Carlander,
1969.
j
Assumed
based
on
Carlander,
1969.
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
3
!

"



	


#
)


$



(

	






























&
Stage
Name
Natural
Mortality
(
per
stage)
a
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryc
Weight
(
lbs)
d
Eggs
1.8
0b
0
0.0000000179e
Larvae
0.498
0b
0
0.00000857e
Age
0
2.93
0b
0
0.012f
Age
1+
0.292
0.292a
0.5
0.128f
Age
2+
0.292
0.292a
1
0.193f
Age
3+
0.292
0.292a
1
0.427f
Age
4+
0.292
0.292a
1
0.651f
Age
5+
0.292
0.292a
1
0.888f
Age
6+
0.292
0.292a
1
0.925f
Age
7+
0.292
0.292a
1
0.972g
Age
8+
0.292
0.292a
1
1.08
f
Age
9+
0.292
0.292a
1
1.26
f
a
Bartell
and
Campbell,
2000.
b
Recreational
species.
Fraction
vulnerable
assumed.
c
Assumed.
d
Weight
calculated
from
length
using
the
formula:
(
1.014*
10­
5)*
Length(
mm)
3.066
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
e
Wang,
1986b.
f
Carlander,
1977.
g
Assumed
based
on
Carlander,
1977.
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
4
!

"



	


#
*


$

 

%



























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisherye
Weight
(
lbs)
g
Eggs
1.73a
0e
0
0.0000000108h
Larvae
0.576a
0e
0
0.00000156h
Age
0
4.62a
0e
0
0.00795i
Age
1+
0.39b
0e
0
0.00992i
Age
2+
0.151c
0e
0
0.032i
Age
3+
0.735d
0.735f
0.5
0.0594i
Age
4+
0.735d
0.735f
1
0.104i
Age
5+
0.735d
0.735f
1
0.189i
Age
6+
0.735d
0.735f
1
0.193i
Age
7+
0.735d
0.735f
1
0.209i
Age
8+
0.735d
0.735f
1
0.352i
Age
9+
0.735d
0.735f
1
0.393i
a
Bartell
and
Campbell,
2000.
b
Froese
and
Pauly,
2001.
c
Calculated
from
survival
(
PSEG,
1999c)
(
Carlander,
1977)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)­(
fishing
mortality).
d
Calculated
from
survival
(
PSEG,
1999c)
(
Carlander,
1977)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival1/
2).
e
Recreational
species.
Fraction
vulnerable
assumed.
f
Calculated
from
survival
(
PSEG,
1999c)
(
Carlander,
1977)
using
the
equation:
(
fishing
mortality)
=
­
LN(
survival1/
2).
g
Weight
calculated
from
length
using
the
formula:
(
4.326*
10­
6)*
Length(
mm)
3.209
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
h
Wang,
1986b.

!

"



	


#
+


$

 







,




-
























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
e
Eggs
2.3a
0
0
0.000000985f
Larvae
2.06b
0
0
0.000375g
Age
0
2.06b
0
0
0.00208g
Age
1+
1c
0
0
0.00585g
Age
2+
1c
0
0
0.0121g
Age
3+
1c
0
0
0.0143f
a
Calculated
from
assumed
survival(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)­
(
fishing
mortality).
b
Calculated
from
estimated
survival
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)­(
fishing
mortality).
c
Froese
and
Pauly,
2001.
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Weight
calculated
from
length
using
the
formula:
(
4.466*
10­
4)*
Length(
mm)
2.34
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
f
Assumed
based
on
Carlander,
1969.
g
Carlander,
1969.
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
5
!

"



	


#
.


$


-


$
 






























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
e
Eggs
2.3a
0
0
0.0000000165f
Larvae
4.61b
0
0
0.0000192f
Age
0
1.39b
0
0
0.00246g
Age
1+
0.446c
0
0
0.0898g
Age
2+
0.446c
0
0
0.172g
Age
3+
0.446c
0
0
0.278g
Age
4+
0.446c
0
0
0.33g
Age
5+
0.446c
0
0
0.57g
Age
6+
0.446c
0
0
0.582g
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
survival
(
PSEG,
1999c)
for
channel
catfish
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
Calculated
from
survival
(
PSEG,
1999c)
(
Carlander,
1969)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Weight
calculated
from
length
using
the
formula:
(
7.491*
10­
6)*
Length(
mm)
3.065
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
f
Assumed
based
on
Carlander,
1969.
g
Carlander,
1969.
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
6
!

"



	


#
/


	







	






























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
f
Eggs
2.3a
0d
0
0.000000408g
Larvae
4.61b
0d
0
0.0000191g
Age
0
1.39b
0d
0
0.00987h
Age
1+
0.41c
0.41e
0.5
0.0554h
Age
2+
0.41c
0.41e
1
0.189h
Age
3+
0.41c
0.41e
1
0.436h
Age
4+
0.41c
0.41e
1
0.71
h
Age
5+
0.41c
0.41e
1
1.22
h
Age
6+
0.41c
0.41e
1
1.55
h
Age
7+
0.41c
0.41e
1
2.27
h
Age
8+
0.41c
0.41e
1
2.66
h
Age
9+
0.41c
0.41e
1
3.41
h
Age
10+
0.41c
0.41e
1
5.59
h
Age
11+
0.41c
0.41e
1
5.81
i
Age
12+
0.41c
0.41e
1
5.92
h
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
survival
(
PSEG,
1999c)
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
Calculated
from
survival(
PSEG,
1999c)
(
Miller,
1966)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival1/
2).
d
(
Salia
et
al.,
1997)
Hake.
e
Calculated
from
survival
(
PSEG,
1999c)
(
Miller,
1966)
using
the
equation:
(
fishing
mortality)
=
­
LN(
survival1/
2).
f
Weight
calculated
from
length
using
the
formula:
(
2.945*
10­
6)*
Length(
mm)
3.133
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
g
(
Wang,
1986b).
h
(
Carlander,
1969).
i
Assumed
based
on
(
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
7
!

"



	


#
0


	






	



























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
e
Eggs
2.3a
0
0
0.000000143f
Larvae
4.61b
0
0
0.0000118f
Age
0
1.39b
0
0
0.0225g
Age
1+
0.26c
0
0
0.79g
Age
2+
0.26c
0
0
1.21g
Age
3+
0.26c
0
0
1.81g
Age
4+
0.26c
0
0
5.13g
Age
5+
0.26c
0
0
5.52h
Age
6+
0.26c
0
0
5.82h
Age
7+
0.26c
0
0
6.76g
Age
8+
0.26c
0
0
8.17g
Age
9+
0.26c
0
0
8.55h
Age
10+
0.26c
0
0
8.94h
Age
11+
0.26c
0
0
9.76h
Age
12+
0.26c
0
0
10.2h
Age
13+
0.26c
0
0
10.6h
Age
14+
0.26c
0
0
11.1h
Age
15+
0.26c
0
0
11.5h
Age
16+
0.26c
0
0
12
h
Age
17+
0.26c
0
0
12.5h
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
survival
(
PSEG,
1999c)
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Weight
calculated
from
length
using
the
formula:
(
1.095*
10­
5)*
Length(
mm)
3.025
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
f
(
Wang,
1986b).
g
(
Carlander,
1969).
h
Assumed
based
on
(
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
8
!

"



	


#
1


2





































&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisherye
Weight
(
lbs)
f
Eggs
2.3a
0
0
0.000000252e
Larvae
2.3b
0
0
0.0016e
Age
0
1.39b
0
0
0.0135g
Age
1+
0.916b
0
0
0.026g
Age
2+
1.1c
0
0
0.0478g
Age
3+
1.1c
0
0
0.106g
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
survival
(
PSEG,
1999c)
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Assumed.
f
Weight
calculated
from
length
using
the
formula:
(
1.144*
10­
4)*
Length(
mm)
2.92
=
Weight(
gm)
(
Fuchs,
1967).
g
(
Trautman,
1981).

!

"



	


#


3







-





4

 

























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)

Eggs
2.3a
0
0
0.0000011e
Larvae
6.13b
0
0
0.00000295b
Age
0
2.3b
0
0
0.0166b
Age
1+
0.31c
0
0
0.05f
Age
2+
0.31c
0
0
0.206f
Age
3+
0.31c
0
0
0.438f
Age
4+
0.31c
0
0
0.638f
Age
5+
0.31c
0
0
0.794f
Age
6+
0.31c
0
0
0.95f
Age
7+
0.31c
0
0
1.09f
Age
8+
0.31c
0
0
1.26f
Age
9+
0.31c
0
0
1.44f
Age
10+
0.31c
0
0
1.6
f
Age
11+
0.31c
0
0
1.78f
Age
12+
0.31c
0
0
2
f
a
Calculated
from
assumed
survival(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
(
Bartell
and
Campbell,
2000).
c
(
Froese
and
Pauly,
2001).
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Assumed
based
on
(
Bartell
and
Campbell,
2000).
f
(
Scott
and
Crossman,
1973).
Average
of
males
and
females.
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
9
!

"



	


#






5

6
6
































&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
c
Fraction
Vulnerable
to
Fisheryc
Weight
(
lbs)
d
Eggs
2.3a
0
0
0.0000000118e
Larvae
2.3b
0
0
0.0000335e
Age
0
3.69b
0
0
0.0276f
Age
1+
0.916b
0
0
0.322f
Age
2+
0.916b
0
0
0.666f
Age
3+
0.916b
0
0
2.92f
Age
4+
0.916b
0
0
3.66f
Age
5+
0.916b
0
0
4.57f
Age
6+
0.916b
0
0
4.72g
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
survival
(
PSEG,
1999c)
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
d
Weight
calculated
from
length
using
the
formula:
(
1.291*
10­
5)*
Length(
mm)
3.071
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
e
(
Wallus
et
al.,
1990).
f
(
Carlander,
1969).
g
Assumed
based
on
(
Carlander,
1969).

!

"



	


#


'


5






































&
Stage
Name
Natural
Mortality
(
per
stage)
a
Fishing
Mortality
(
per
stage)
b
Fraction
Vulnerable
to
Fisheryb
Weight
(
lbs)
c
Eggs
2.3
0
0
0.0000000308d
Larvae
2.3
0
0
0.0000037d
Age
0
2.99
0
0
0.0267e
Age
1+
0.548
0
0
0.0521e
Age
2+
0.548
0
0
0.18e
Age
3+
0.548
0
0
0.493e
Age
4+
0.548
0
0
0.653e
Age
5+
0.548
0
0
0.916e
Age
6+
0.548
0
0
2.78e
Age
7+
0.548
0
0
3.07e
a
(
Bartell
and
Campbell,
2000).
Shorthead
redhorse.
b
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
c
Weight
calculated
from
length
using
the
formula:
(
1.398*
10­
5)*
Length(
mm)
2.975=
Weight(
gm)
(
Froese
and
Pauly,
2001).
d
Assumed
based
on
Carlander
(
1969).
e
(
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
10
!

"



	


#


)




%





























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
e
Eggs
2.3a
0
0
0.00000000309f
Larvae
1.95b
0
0
0.000276d
Age
0
1.95b
0
0
0.00345f
Age
1+
0.7c
0
0
0.0128f
Age
2+
0.7c
0
0
0.0274f
Age
3+
0.7c
0
0
0.0443f
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
estimated
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Weight
calculated
from
length
using
the
formula:
(
5.240*
10­
7)*
Length(
mm)
3.641
=
Weight(
gm)
(
Carlander,
1997).
f
(
Carlander,
1997).

!

"



	


#


*





%





 





























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
f
Eggs
1.71a
0d
0
0.00000000736g
Larvae
0.687b
0d
0
0.000000994g
Age
0+
0.687b
0d
0
0.000878h
Age
1+
1.61a
0d
0
0.00666h
Age
2+
1.61a
0d
0
0.0271h
Age
3+
1.5c
1.5e
0.5
0.0593h
Age
4+
1.5c
1.5e
1
0.0754h
Age
5+
1.5c
1.5e
1
0.142h
Age
6+
1.5c
1.5e
1
0.18
h
Age
7+
1.5c
1.5e
1
0.214h
Age
8+
1.5c
1.5e
1
0.232h
a
Calculated
from
survival
for
pumpkinseed
(
Carlander,
1977)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­(
fishing
mortality).
b
Calculated
from
estimated
survival(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
Calculated
from
survival
for
pumpkinseed
(
Carlander,
1977)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival1/
2).
d
Assumed.
e
Calculated
from
survival
for
pumpkinseed
(
Carlander,
1977)
using
the
equation:
(
fishing
mortality)
=
­
LN(
survival1/
2).
f
Weight
calculated
from
length
using
the
formula:
(
3.337*
10­
6)*
Length(
mm)
3.262
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
Pumpkinseed.
g
(
Wang,
1986b).
h
(
Carlander,
1977).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
11
!

"



	


#


+


,
 

(



 

%

























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryf
Weight
(
lbs)
g
Eggs
1.08a
0e
0
0.000000205h
Larvae
5.49b
0e
0
0.0133i
Age
0
5.49b
0e
0
0.0451h
Age
1+
0.15c
0e
0
0.365h
Age
2+
0.15c
0e
0
1.1
h
Age
3+
0.15c
0e
0
1.53
h
Age
4+
0.15c
0e
0
2.72
h
Age
5+
0.15c
0e
0
6.19
h
Age
6+
0.15c
0e
0
7.02
h
Age
7+
0.15c
0e
0
8.92
h
Age
8+
0.15c
0e
0
12.3h
Age
9+
0.15c
0e
0
13.9h
Age
10+
0.075d
0.075d
0.5
16.6h
Age
11+
0.075d
0.075d
1
19
h
Age
12+
0.075d
0.075d
1
24.2h
Age
13+
0.075d
0.075d
1
25.3h
Age
14+
0.075d
0.075d
1
30
h
Age
15+
0.075d
0.075d
1
32.4h
Age
16+
0.075d
0.075d
1
34.3h
Age
17+
0.075d
0.075d
1
45.6h
Age
18+
0.075d
0.075d
1
45.8i
Age
19+
0.075d
0.075d
1
47.7h
Age
20+
0.075d
0.075d
1
48.8i
Age
21+
0.075d
0.075d
1
48.9i
Age
22+
0.075d
0.075d
1
49
i
Age
23+
0.075d
0.075d
1
49.1i
Age
24+
0.075d
0.075d
1
49.2i
Age
25+
0.075d
0.075d
1
49.3i
Age
26+
0.075d
0.075d
1
49.4i
Age
27+
0.075d
0.075d
1
49.4i
a
Calculated
from
survival
(
Carlander,
1969)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
estimated
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
d
(
Froese
and
Pauly,
2001),
assuming
half
of
mortality
was
natural
and
half
was
fishing.
e
Assumed.
f
(
Pennsylvania,
1999).
g
Weight
calculated
from
length
using
the
formula:
(
5.590*
10­
6)*
Length(
mm)
3.016
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
h
(
Carlander,
1969).
i
Assumed
based
on
(
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
12
!

"



	


#


.




































&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisherye
Weight
(
lbs)
f
Eggs
2.3a
0e
0
0.00000361g
Larvae
3.23b
0e
0
0.0000816h
Age
0
3.23b
0e
0
0.0578h
Age
1+
0.57c
0e
0
0.453g
Age
2+
0.285d
0.285d
0.5
7.1g
Age
3+
0.285d
0.285d
1
16.3g
Age
4+
0.285d
0.285d
1
27.4g
Age
5+
0.285d
0.285d
1
31.6g
Age
6+
0.285d
0.285d
1
37.3g
Age
7+
0.285d
0.285d
1
41.6g
Age
8+
0.285d
0.285d
1
43.7g
Age
9+
0.285d
0.285d
1
49.2g
Age
10+
0.285d
0.285d
1
51.9g
Age
11+
0.285d
0.285d
1
54.6g
Age
12+
0.285d
0.285d
1
60.6g
Age
13+
0.285d
0.285d
1
63.5g
Age
14+
0.285d
0.285d
1
68.1g
Age
15+
0.285d
0.285d
1
72.7g
Age
16+
0.285d
0.285d
1
75.5h
Age
17+
0.285d
0.285d
1
80.8h
Age
18+
0.285d
0.285d
1
82.6h
Age
19+
0.285d
0.285d
1
85.4g
Age
20+
0.285d
0.285d
1
87.9g
Age
21+
0.285d
0.285d
1
96.2h
Age
22+
0.285d
0.285d
1
102g
a
Calculated
from
assumed
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
estimated
survival
(
PSEG,
1999c)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
Mississippi
paddlefish.
d
(
Froese
and
Pauly,
2001).
Mississippi
paddlefish.
Assumed
half
of
mortality
was
natural
and
half
was
fishing.
e
Assumed.
f
Weight
calculated
from
length
using
the
formula:
(
5.226*
10­
5)*
Length(
mm)
2.85
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
g
(
Carlander,
1969).
h
Assumed
based
on
(
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
13
!

"



	


#


/






"

-






























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryc
Weight
(
lbs)
e
Eggs
3.32a
0c
0
0.0000000115f
Larvae
2.65a
0c
0
0.00000233f
Age
1+
0.72b
0c
0
0.0195g
Age
2+
0.72b
0c
0
0.041h
Age
3+
0.72b
0d
0
0.177h
Age
4+
0.72b
0d
0
0.338i
Age
5+
0.72b
0d
0
0.537i
Age
6+
0.72b
0d
0
0.597j
a
Calculated
from
survival
(
Carlander,
1969)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
(
Froese
and
Pauly,
2001).
c
Assumed.
d
(
Stone
&
Webster
Engineering
Corporation,
1977).
e
Weight
calculated
from
length
using
the
formula:
(
5.23*
10­
6)*
Length(
mm)
3.114
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
f
(
Able
and
Fahay,
1998).
g
Assumed
based
on
(
Able
and
Fahay,
1998)
and
(
Scott
and
Scott,
1988).
h
(
Scott
and
Scott,
1988).
i
Assumed
based
on
(
Scott
and
Scott,
1988).
j
Assumed
to
be
5.6
cm
less
than
maximum
reported
length
(
Froese
and
Pauly,
2001).

!

"



	


#


0








	




 

(


























&
Stage
Name
Natural
Mortality
(
per
stage)
a
Fishing
Mortality
(
per
stage)
b
Fraction
Vulnerable
to
Fisheryb
Weight
(
lbs)
c
Eggs
2.05
0
0
0.0000000135d
Larvae
2.56
0
0
0.00000198d
Age
0+
2.3
0
0
0.000145e
Age
1+
0.548
0
0
0.0447e
Age
2+
0.548
0
0
0.249e
Age
3+
0.548
0
0
0.305e
Age
4+
0.548
0
0
0.609e
Age
5+
0.548
0
0
0.823e
Age
6+
0.548
0
0
0.929e
a
(
Bartell
and
Campbell,
2000).
b
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
c
Weight
calculated
from
length
using
the
formula:
(
6.13*
10­
6)*
Length(
mm)
3.099
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
d
Assumed
based
on
Carlander
(
1969).
e
(
Carlander,
1969).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
14
!

"



	


#


1




 
%


























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
c
Fraction
Vulnerable
to
Fisheryc
Weight
(
lbs)
d
Eggs
1.05a
0
0
0.0000000179c
Larvae
3.55b
0
0
0.000163e
Age
0+
1.62b
0
0
0.0341f
Age
1+
0.23b
0.05
0.5
0.505f
Age
2+
0.23b
0.05
1
1.03f
Age
3+
0.23b
0.05
1
1.53f
Age
4+
0.23b
0.05
1
2.19f
Age
5+
0.23b
0.05
1
2.27f
Age
6+
0.23b
0.05
1
3.82f
Age
7+
0.23b
0.05
1
4.65f
Age
8+
0.23b
0.05
1
4.8f
a
Calculated
from
survival
for
walleye
(
Carlander,
1997)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
(
Bartell
and
Campbell,
2000).
c
Recreational
species.
Fraction
vulnerable
assumed.
d
Weight
calculated
from
length
using
the
formula:
(
8.137*
10­
6)*
Length(
mm)
3.058
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
e
(
Carlander,
1997).
Walleye.
f
(
Carlander,
1997).

!

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
	


#
'
3



(


7


(







%
























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
e
Eggs
2.3a
0
0
0.00000242f
Larvae
4.25b
0
0
0.000381f
Age
0+
4.25b
0
0
0.0572f
Age
1+
0.7c
0
0
0.301g
Age
2+
0.7c
0
0
0.833g
Age
3+
0.7c
0
0
1.74h
a
Calculated
from
assumed
survival
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
Calculated
from
estimated
survival
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
Alewife.
d
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
e
Assumed.
e
Weight
calculated
from
length
using
the
formula:
(
7.565*
10­
5)*
Length(
mm)
2.690
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
f
(
Wallus
et
al.,
1990).
g
Assumed
based
on
(
Wallus
et
al.,
1990)
and
(
Trautman,
1981).
h
(
Trautman,
1981).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
15
!

"



	


#
'











 



$



























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryd
Weight
(
lbs)
f
Eggs
1.9a
0d
0
0.000000331g
Larvae
2.7b
0d
0
0.0000198g
Age
0+
0.446a
0d
0
0.0169c
Age
1+
0.86c
0.23c
0.5
0.202c
Age
2+
1.17c
0.322e
1
0.518c
Age
3+
0.755c
0.208e
1
0.733c
Age
4+
1.05c
0.288e
1
1.04
c
Age
5+
0.867c
0.238e
1
1.44
c
Age
6+
0.867c
0.238e
1
2.24
c
Age
7+
0.867c
0.238e
1
2.56
h
Age
8+
0.867c
0.238e
1
2.92
h
Age
9+
0.867c
0.238e
1
3.3
c
a
Calculated
from
survival
(
Carlander,
1977)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
(
Bartell
and
Campbell,
2000).
c
(
Carlander,
1977).
d
Recreational
species.
Fraction
vulnerable
assumed.
e
(
Carlander,
1977),
assuming
that
27.5%
of
natural
mortality
was
fishing
mortality.
f
Weight
calculated
from
length
using
the
formula:
(
2.494*
10­
5)*
Length(
mm)
2.917
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
g
(
Wang,
1986b).
h
Assumed
based
on
(
Carlander,
1977).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
16
!

"



	


#
'
'










$



























&
Stage
Name
Natural
Mortality
(
per
stage)
a
Fishing
Mortality
(
per
stage)
a
Fraction
Vulnerable
to
Fisherya
Weight
(
lbs)
b
Eggs
1.39
0
0
0.000022b
Yolksac
larvae
2.22
0
0
0.097c
Post­
yolksac
larvae
5.11
0
0
0.194c
Juvenile
1
2.28
0
0
0.291c
Juvenile
2
1
0
0
0.388c
Age
1+
1.1
0
0
0.485a
Age
2+
0.15
0.31
0.06
2.06a
Age
3+
0.15
0.31
0.2
3.31a
Age
4+
0.15
0.31
0.63
4.93a
Age
5+
0.15
0.31
0.94
6.5a
Age
6+
0.15
0.31
1
8.58a
Age
7+
0.15
0.31
0.9
12.3a
Age
8+
0.15
0.31
0.9
14.3a
Age
9+
0.15
0.31
0.9
16.1a
Age
10+
0.15
0.31
0.9
18.8a
Age
11+
0.15
0.31
0.9
19.6a
Age
12+
0.15
0.31
0.9
22.4a
Age
13+
0.15
0.31
0.9
27a
Age
14+
0.15
0.31
0.9
34.6a
Age
15+
0.15
0.31
0.9
41.5a
a
(
PSEG,
1999c).
b
Assumed.
c
Assumed
based
on
(
PSEG,
1999c).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
17
!

"



	


#
'
)



 

(


























&
Stage
Name
Natural
Mortality
(
per
stage)
a
Fishing
Mortality
(
per
stage)
b
Fraction
Vulnerable
to
Fisheryb
Weight
(
lbs)
c
Eggs
2.05
0
0
0.0000000135d
Larvae
2.56
0
0
0.00000198d
Age
0+
2.3
0
0
0.000145e
Age
1+
0.548
0
0
0.0447e
Age
2+
0.548
0
0
0.249e
Age
3+
0.548
0
0
0.305e
Age
4+
0.548
0
0
0.609e
Age
5+
0.548
0
0
0.823e
Age
6+
0.548
0
0
0.929e
a
(
Bartell
and
Campbell,
2000).
b
Not
a
commercial
or
recreational
species,
thus
no
fishing
mortality.
c
Weight
calculated
from
length
using
the
formula:
(
6.130*
10­
5)*
Length(
mm)
3.099
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
River
carpsucker.
d
Assumed
based
on
(
Carlander,
1969).
e
(
Carlander,
1969).

!

"



	


#
'
*


8






























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryc
Weight
(
lbs)
e
Eggs
1.05a
0c
0
0.00000000506f
Larvae
3.55b
0c
0
0.0000768g
Age
0+
1.93b
0c
0
0.03
g
Age
1+
0.0474b
0.6d
0.5
0.328g
Age
2+
0.0474b
0.6d
1
0.907g
Age
3+
0.0474b
0.6d
1
1.77
g
Age
4+
0.0474b
0.6d
1
2.35
g
Age
5+
0.0474b
0.6d
1
3.37
g
Age
6+
0.0474b
0.6d
1
3.97
g
Age
7+
0.0474b
0.6d
1
4.66
f
Age
8+
0.0474b
0.6d
1
5.58
g
a
Calculated
from
survival
(
Carlander,
1997)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
b
(
Bartell
and
Campbell,
2000).
c
Recreational
species.
Fraction
vulnerable
assumed.
d
(
McDermot
and
Rose,
2000).
e
Weight
calculated
from
length
using
the
formula:
(
2.296*
10­
6)*
Length(
mm)
3.23
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
f
Assumed
based
on
(
Carlander,
1997).
g
(
Carlander,
1997).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
18
!

"



	


#
'
+


8





$



























&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
d
Fraction
Vulnerable
to
Fisherye
Weight
(
lbs)

Eggs
2.3a
0
0
0.0000000266f
Larvae
4.61b
0
0
0.00000174g
Age
0+
1.39b
0
0
0.174h
Age
1+
0.42c
0
0
0.467h
Age
2+
0.42c
0.7
0.5
0.644h
Age
3+
0.42c
0.7
1
1.02h
Age
4+
0.42c
0.7
1
1.16h
Age
5+
0.42c
0.7
1
1.26h
Age
6+
0.42c
0.7
1
1.66h
Age
7+
0.42c
0.7
1
1.68i
a
Calculated
from
assumed
survival
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)­(
fishing
mortality).
b
Calculated
from
survival
(
Geo­
Marine
Inc.,
1978)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)­
(
fishing
mortality).
c
(
Froese
and
Pauly,
2001).
d
(
McDermot
and
Rose,
2000).
e
Recreational
species.
Fraction
vulnerable
assumed.
f
Calculated
from
assumed
length
of
1
mm
using
the
formula:
(
1.206*
10­
5)*
Length(
mm)
3.132
=
Weight(
gm)
(
Van
Oosten,
1942).
g
Calculated
from
length
of
3.8
mm
(
Carlander,
1997)
using
the
formula:
(
1.206*
10­
5)*
Length(
mm)
3.132
=
Weight(
gm)
(
Van
Oosten,
1942).
h
(
Carlander,
1997).
i
Assumed
based
on
(
Carlander,
1997).
†





E


&
D
V
H

6
W
X
G
L
H
V


3
D
U
W

&


7
K
H

2
K
L
R

5
L
Y
H
U

$
S
S
H
Q
G
L
[

&

App.
C1­
19
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#
'
.

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9
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-
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







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
&
Stage
Name
Natural
Mortality
(
per
stage)
Fishing
Mortality
(
per
stage)
Fraction
Vulnerable
to
Fisheryg
Weight
(
lbs)
h
Eggs
2.75a
0e
0
0.0000000274i
Larvae
1.79b
0e
0
0.00171j
Age
0+
1.79b
0e
0
0.01
j
Age
1+
2.41c
0e
0
0.051j
Age
2+
1.35c
0e
0
0.137j
Age
3+
1.2c
0e
0
0.23
j
Age
4+
1.24c
0e
0
0.347j
Age
5+
1.31c
0e
0
0.46
j
Age
6+
1.77c
0e
0
0.617j
Age
7+
1.83d
0.7f
0.5
0.763j
Age
8+
1.83d
0.7f
1
0.92
j
Age
9+
1.83d
0.7f
1
1.15
k
Age
10+
1.83d
0.7f
1
1.17
j
a
(
PSEG,
1999c).
White
perch.
b
Calculated
from
estimated
survival
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
c
Calculated
from
survival
(
Carlander,
1997)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality).
d
Calculated
from
survival
(
Carlander,
1997)
using
the
equation:
(
natural
mortality)
=
­
LN(
survival)
­
(
fishing
mortality),
and
subtracting
known
fishing
mortality
(
McDermot
and
Rose,
2000).
e
Assumed
based
on
landings
data.
See
Chapter
C6.
f
(
McDermot
and
Rose,
2000).
g
Recreational
species.
Fraction
vulnerable
assumed.
h
Weight
calculated
from
length
using
the
formula:
(
1.710*
10­
5)*
Length(
mm)
3.015
=
Weight(
gm)
(
Froese
and
Pauly,
2001).
i
(
Wang
and
Kernehan,
1979).
j
(
Carlander,
1997).
k
Assumed
based
on
(
Carlander,
1997).
